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mechanoreceptors that are sensitive to hair movement, skin pressure, or noxious stimuli There are also descending bers in the posterior columns, including bers from cells in the dorsal column nuclei The posterior columns contain a portion of the bers for the sense of touch as well as the bers mediating the senses of touchpressure, vibration, direction of movement and position of joints, and stereoesthesia recognition of surface texture, shape, numbers and gures written on the skin, and two-point discrimination all of which depend on patterns of touch-pressure (see Fig 8-2) The nerve cells of the nuclei gracilis and cuneatus and accessory cuneate nuclei give rise to a secondary afferent path, which crosses the midline in the medulla and ascends as the medial lemniscus to the posterior thalamus However, the ber pathways in the posterior columns are not the sole mediators of proprioception in the spinal cord [see Posterior (Dorsal) Column Syndrome, further on] In addition to these well-de ned posterior column pathways, there are cells in the reticular part of the dorsal column nuclei that receive secondary ascending bers from the dorsal horns of the spinal cord and from ascending bers in the posterolateral columns These dorsal column bers project to brainstem nuclei, cerebellum, and

S1 L5 Figure 9-2 Distribution of the sensory spinal roots on the surface of the body (Reproduced by permission from Sinclair)

FIGURE 5-11

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cord, terminating in the gracile and cuneate nuclei The central axons of the primary sensory neurons are joined in the posterior columns by other secondary neurons whose cell bodies lie in the posterior horns of the spinal cord (see below) The bers in the posterior columns are displaced medially as new bers from each successively higher root are added, thereby creating somatotopic laminations (see Fig 8-3) Of the long ascending posterior column bers, which are activated by mechanical stimuli of skin and subcutaneous tissues and by movement of joints, only about 25 percent (from the lumbar region) reach the gracile nuclei at the upper cervical cord The rest off collaterals to or terminate in the dorsal horns of the spinal cord, at least in the cat (Davidoff) An estimated 20 percent of ascending bers originate from cells in Rexed layers IV and V of the posterior horns (see Fig 8-1) and convey impulses from low-threshold

A number of different symptoms are associated with mice, and they provide a common source of computer problems Fortunately, most procedures to resolve a mouse problem are quite simple Let s look at two common mouse-related problems

Figure 9-3 Dermatomes of the upper and lower extremities, outlined by the pattern of sensory loss following lesions of single nerve roots (Reproduced by permission from Keegan and Garrett)

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